Zima Y., Fedorenko V. Distribution of snakes of the vipera family Viperidae in Kazakhstan and modeling of their potential ranges // Principy èkologii. 2022. № 1. P. 16‒46. DOI: 10.15393/j1.art.2022.12424

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Distribution of snakes of the vipera family Viperidae in Kazakhstan and modeling of their potential ranges

Abduraupov T. V. Pestov M. V. Nuridzhanov A. S. Caruk O. I. Kirshey T. Preliminary review of modern herpetofauna of the southern part of Karakalpak Ustyurt, Biologicheskie i strukturno-funkcional'nye osnovy izucheniya i sohraneniya bioraznoobraziya Uzbekistana: Materialy Respublikanskoy nauchnoy konferencii, posvyaschennoy 80-letiyu professora A. A. Butnik (Tashkent, 8−9 sentyabrya 2015 g.). Tashkent, 2015. P. 17−22.

Ahmedenov K. M. Abuova R. S. Distribution of the eastern steppe viper Vipera renardi (Reptilia, Viperidae) in the West Kazakhstan region, Karantinnye i zoonoznye infekcii v Kazahstane. Almaty, 2018. No. 1−2 (36−37). P. 110−117.

Ahmedenov K. M. Petrischev V. P. Golovachev I. V. Bakiev A. G. Gorelov R. A. Kalmykova O. G. Maykanov N. S. Indersky salt-dome landscape - a reserved pearl of Western Kazakhstan: Monograph. Ural'sk: Zap, Kazahst. agr, tehn. un-t im. Zhangir hana, 2017. 142 p.

Alekperov A. M. Amphibians and reptiles of Azerbaijan. Baku: ELM, 1978. 264 p.

Alekperov A. M. Poisonous snakes of Azerbaijan and issues of their protection, Yadovitye zhivotnye Sredney Azii i ih yady: Materialy Sredneaziatskoy konferencii (1−3 oktyabrya 1968 g.). Tashkent, 1970. P. 14−28.

Alenicin V. D. Reptiles of the islands and shores of the Aral Sea. SPb., 1876. 64 p. (Trudy Aralo-Kaspiyskoy ekspedicii. Vyp. 3).

Aliev S. A. Amphibians and reptiles of Tajikistan. Dushanbe: Donish, 1979. 147 p.

Aliev T. To the study of some poisonous snakes of Azerbaijan, Voprosy gerpetologii. L.: Nauka, 1973. P. 6−8.

Amphibians and reptiles of the Orenburg region and their protection. Materials for the Red Book of the Orenburg region. Ekaterinburg: UrO RAN, 1995. 45 p.

Anan'eva N. B. Munhbayar X. Orlov N. L. Orlova V. F. Semenov D. V. Terbish H. Amphibians and reptiles of Mongolia. Reptiles. M.: KMK Ltd, 1997. 416 p.

Anan'eva N. B. Orlov N. L. Halikov R. G. Darevskiy I. S. Ryabov S. A. Barabanov A. V. Atlas of reptiles of Northern Eurasia (taxonomic diversity, geographical distribution and conservation status). SPb.: Zoologicheskiy institut, 2004. 232 p.

Andrushko A. M. Reptiles of the Kazakh Highlands and their economic importance, Uchenye zapiski LGU. Ser. biol. 1955. T. 181, No. 38. P. 19−43.

Andryuschenko A. V. To the question of the species composition of the herpetofauna of the Kostanay region, Biologicheskoe raznoobrazie aziatskih stepey: Materialy mezhdunar. nauch. konf. Kostanay, 2007. P. 7−9.

Antipin V. M. Outline of terrestrial vertebrates of the Karatau Ridge, Byulleten' Moskovskogo obschestva ispytateley prirody. Otd. biol. 1955. T. 60, No. 1. P. 33−38.

Araujo M. B., Pearson R. G., Thuiller W., Erhard M. Validation of species–climate impact models under climate change, Global Change Biology. 2005. No. 11. P. 1504−1513.

Asadi A., Montgelard C., Nazarizadeh M., Moghaddasi A., Fatemizadeh F., Simonov E., Kami H. G., Kaboli M. Evolutionary history and postglacial colonization of an Asian pit viper (Gloydius halys caucasicus) into Transcaucasia revealed by phylogenetic and phylogeographic analyses, Scientific Reports. 2019. Vol. 9, No. 1224. P. 1−16.

Ataev Ch. A. Reptiles of the mountains of Turkmenistan. Ashhabad: Ylym, 1985. 344 p.

Bakiev A. G. Fayzulin A. I. Materials for the cadastre of amphibians and reptiles of the Middle Volga basin. N. Novgorod, 2002. 132 p.

Bakiev A. G. Garanin V. I. Gelashvili D. B. Gorelov R. A. Doronin I. V. Zayceva O. V. Zinenko A. I. Makarova T. N. Pavlov A. V. Petrova I. V. Ratnikov V. Yu. Starkov V. G. Shiryaeva I. V. Yusupov R. H. Yakovleva T. I. Vipers (Reptilia: Serpentes: Viperidae: Vipera) of the Volga basin. Part 1. Tol'yatti: OOO «Kassandra», 2015. 234 p.

Bakiev A. G. Garanin V. I. Litvinov N. A. Pavlov A. V. Ratnikov V. Yu. Snakes of the Volga-Kama Territory. Samara: Izd-vo Samarskogo nauchnogo centra RAN, 2004. 192 p.

Bakiev A. G. Malenev A. L. Zayceva O. V. Shurshina I. V. Snakes of the Samara region. Tol'yatti: OOO «Kassandra», 2009. 170 p.

Bannikov A. G. Darevskiy I. S. Ischenko V. G. Rustamov A. K. Scherbak N. N. Key to Amphibians and Reptiles of the USSR Fauna. M.: Prosveschenie, 1977. 416 p.

Bartenev A. N. To the issue of zoogeography of the north in the Karaganda region, Uchenye zapiski KazGU. T. 1. Biologiya. Alma-Ata, 1938. P. 67−80.

Bazarbekov K. U. Lyahov O. V. Fauna of the Pavlodar Irtysh region (vertebrates). Pavlodar: Pavlodarskiy gop. un-t im. P. Toraygyrova, 2004. 336 p.

Bebutov A. M. Observations on reptiles of Central Kazakhstan on the route of the Julek village − Atbasar city, Trudy instituta geografii AN SSSR. 1953. No. 54. P. 423−434.

Bekenov A. B. Erzhanov N. T. Kapitonov V. I. Slavchenko N. P. Berber A. P. Isenov H. A. Abukenova B. C. Rare and endangered animals of the Kazakh uplands. Pavlodar: Izd-vo PGU im. P. Toraygyrova, 2005. 363 p.

Berdibaeva Zh. Sh. Materials on the ecology of the Halys pit viper, Biologicheskie nauki: Sb. statey. Vyp. 1. Alma-Ata, 1974. P. 67−69.

Berdibaeva Zh. Sh. Seasonal changes in the state of developing eggs in reptiles of the Upper Irtysh region, Issledovaniya i metodicheskie raboty po biologicheskim naukam: Materialy zonal'nogo soveschaniya pri Ust'-Kam. ped. in-te. Vyp. 1. Alma-Ata, 1966. P. 16−19.

Berdibaeva Zh. Sh. To the biology of the common adder, Fauna i ekologiya zhivotnyh Kazahstana. Alma-Ata, 1981. P. 31−33.

Berezovikov N. N. Hrokov V. V. Brushko Z. K. Mitrofanov I. V. Bragin B. I. Nilov V. I. Kornelyuk A. I. Shaymardanov R. T. Murzov V. N. The impact of the development of the Karachaganak oil and gas condensate field on the fauna of Western Kazakhstan, Selevinia. 2003. T. 1, No. 4. P. 123−137.

Berezovikov N. N. About the case of the death of a widgeon (Anas penelope) from a viper bite, Selevinia. 1994. T. 2, No. 4. C. 81.

Bobrov V. V. Materials on the fauna and population of reptiles of the North-Western Caspian, Voprosy gerpetologii. L.: Nauka, 1985. P. 29−30.

Bogdanov O. P. SR. T. Fauna of the Uzbek SSR. Vol. 1. Amphibians and reptiles. Tashkent: Izd-vo AN UzSSR, 1960. 260 p.

Bogdanov O. P. Animals of Uzbekistan (vertebrates). Tashkent, 1961. 316 p.

Bogdanov O. P. Ecology of reptiles in Central Asia. Tashkent: FAN, 1965. 260 p.

Bogdanov O. P. Feeding of the Halys pit viper (Ancistrodon halys) in Central Asia, Zoologicheskiy zhurnal. 1970. T. 49, No. 12. P. 1851−1856.

Bondarenko D. A. Peregoncev E. A. Communities of reptiles of Karakalpak Ustyurt (Uzbekistan), Sovremennaya gerpetologiya. 2018. T. 18, No. 1/2. P. 13−26.

Borkin L. Ya. Gannibal B. K. Golubev A. V. The roads of Peter Simon Pallas (in the west of Kazakhstan). SPb.; Ural'sk: Evraziyskiy soyuz uchenyh, 2014. 312 p.

Borkin L. Ya. Il'yashenko V. Yu. Herpetofauna of the Zeya Reserve and the surroundings of the city of Zeya (Amur Region), Voprosy gerpetologii. L.: Nauka, 1981. P. 21−22.

Borkin L. Ya. Litvinchuk S. N. Herpetological research in the west of Kazakhstan: P. S. Pallas and modernity, Priroda zapadnogo Kazahstana i Petr Simon Pallas (polevye issledovaniya 2012 goda). SPb.: Evropeyskiy Dom, 2015. P. 53−79.

Bragina T. M. Bragin E. A. Natural conditions and fauna of the Altyn Dala state nature reserve. Kostanay: TOO «Kostanaypoligrafiya», 2017. 236 p.

Bragina T. M. Bragin E. A. The most important wetlands of Northern Kazakhstan (within the Kostanay and western part of the North Kazakhstan regions). M.: Russkiy universitet, 2002. 156 p.

Bragina T. M. Valyaeva E. A. Solov'ev A. I. Species composition, distribution features and morphometric indicators of some representatives of the batrachofauna and herpetofauna of the Kostanay region, Nauchno-metodicheskiy zhurnal. 2011. No. 2. Vyp. 22. P. 57−59.

Brem A. Life of animals. reptiles. M.: OOO «Firma «Izdatel'stvo AST», 2000. 664 p.

Brushko Z. K. New data on the distribution of reptiles in Kazakhstan, Izvestiya AN KazSSR. Ser. biol. 1983. No. 2. C. 35−38.

Bulanov P. A. Poisonous snakes of Kazakhstan, properties of their venom and anti-venom sera and vaccines, Trudy Alma-Atinskogo vetzooinstituta. 1948. T. 4. P. 158−170.

Caruk O. I. New data on the distribution of some species of amphibians and reptiles in South-Eastern Kazakhstan, Voprosy gerpetologii. L.: Nauka, 1981. P. 143−144.

Chernov S. A. SR. Fauna of the Tajik SSR. Reptiles. Vol. 18. Dushanbe, 1959. 204 p.

Chernov S. A. Ecological and faunistic review of reptiles in the south of the Volga-Ural interfluve, Trudy Zoologicheskogo instituta AN SSSR. 1954. T. 16. P. 137−159.

Chernov S. A. Materials on the herpetofauna of the Kazakh highlands, the northern coast of Balkhash and the Kan Tau mountains, Izvestiya Akademii nauk Kazahskoy SSR. Ser. zool. 1947. No. 6. P. 120−126.

Conant R. Agkistrodon in Europe (Reptilia: Serpentes: Viperidae), Salamandra. 1982. Vol. 18, No. 3/4. P. 191−195.

David P., Vogel G. Terralog: Venomous Snakes of Europe, Northern, Central and Western Asia, vol. 16. Frankfurt am Main: Edition Chimaira, 2010. 160 p.

Davletbakov A. T. Kustareva L. A. Mil'ko D. A. Ostaschenko A. N. Sagymbaev S. S. Toropov S. A. Trotchenko N. V. Inventory of the genetic fund of Kyrgyzstan: V. 4: Type Chordata – Chordates. Bishkek, 2015. 128 p.

Debelo P. V. Chibilev A. A. Amphibians and reptiles of the Ural-Caspian region. Series: Natural diversity of the Ural-Caspian region. Vol. 3.. Ekaterinburg: RIO UrO RAN, 2013. 400 p.

Dinesman L. T. Kaleckaya M. L. Quantitative methods for amphibians and reptiles, Metody ucheta chislennosti i geograficheskogo raspredeleniya nazemnyh pozvonochnyh. M.: Izd-vo AN SSSR, 1952. P. 329−341.

Dinesman L. T. Amphibians and reptiles of the southeast of the Turgai table country and the Northern Aral Sea region, Trudy Instituta geografii AN SSSR. 1953. No. 54. P. 383−422.

Docenko I. B. Catalog of the сollections of the Zoological Museum of the NNPM NAS of Ukraine. Snakes. Kiev: Zoomuzey NNPM NAN Ukrainy, 2003. 86 p.

Duysebaeva T. N. Chirikova M. A. Zima Yu. A. Belyalov O. V. Kovalenko A. V. New data on the distribution of amphibians and reptiles in Kazakhstan: an overview of the first decade of the 21st century, Gerpetologicheskie issledovaniya v Kazahstane i sopredel'nyh stranah. Almaty, 2010. P. 84−99.

Duysebaeva T. N. Amphibians and reptiles of the Markakol basin, Selevinia. 2002. No. 1-4. P. 73−86.

Duysebaeva T. N. Brief review of recent changes in systematic list of amphibians and reptiles of Kazakhstan, Gerpetologicheskie issledovaniya v Kazahstane i sopredel'nyh stranah. Almaty, 2010. P. 37−52.

Duysebaeva T. N. New finds of amphibians and reptiles in the Aral Sea region and adjacent regions of Kazakhstan. Part 2. Snakes (Reptilia: Squamata: Serpentas), Selevinia. 2005. P. 49−56.

Elith J., Graham C. H., Anderson R. P., Dudik M., Ferrier S., Guisan A., Hijmans R. J., Huettmann F., Leathwick J. R., Lehmann A., Li J., Lohmann L. G., Loiselle B. A., Manion G., Moritz C., Nakamura M., Nakazawa Y., Overton J. McC., Peterson A. T., Phillips S. J., Richardson K. S., Scachetti-Pereira R., Schapire R. E., Soberon J., Williams S., Wisz M. S. and Zimmermann N. E. Novel methods improve prediction of species’ distributions from occurrence data, Ecography. 2006. No. 29. P. 129−151.

Elith J., Phillips S. J., Hastie T., Dudik M., Chee Y. E., Yates C. J. A statistical explanation of MaxEnt for ecologists, Diversity and Distributions. 2011. Vol. 17. P. 43−57.

Elpat'evskiy V. S. Reptiles and amphibians collected by the Balkhash expedition in 1903 on the banks of lake Balkhash and the Ili river, Izvestiya Turkestanskogo otdeleniya IRGO. T. 4. Nauchnye rezul'taty Aral'skoy ekspedicii. No. 7. Tashkent, 1907. P. 49−59.

Eremchenko V. K. Panfilov A. M. Carinenko E. I. New data on the distribution of reptiles in the Tien Shan and North-Eastern Gissar-Alay, Vestnik Kyrgyzskogo gosudarstvennogo nacional'nogo universiteta. Biologiya. 2000. Ser. 3, No. 2. P. 25−28.

Felding A. H., Bell J. F. A review of methods for the assessment of prediction errors in conservation presence/absence models, Environmental Conservation. 1997. Vol. 24, No. 1. P. 38−49.

Fomina M. I. Ecology of the steppe viper in the Chu-Ili interfluve and snake nursery: Abstract of PhD thesis. Tashkent, 1966. 21 p.

Freitas I., Ursenbacher S., Mebert K., Zinenko O., Schweiger S., Wüster W., Brito J.C., Crnobrnja-Isailovi´c J., Halpern B., Fahd S., Santos X., Pleguezuelos J. M., Joger U., Orlov N., Mizsei E., Lourdais O., Zuffi M. A. L., Strugariu A., Zamfirescu S. R., Martínez-Solano Í., Velo-Antón G., Kaliontzopoulou A., Martínez-Freiría F. Evaluating taxonomic inflation: towards evidence-based species delimitation in Eurasian vipers (Serpentes: Viperinae), Amphibia-Reptilia. 2020. Vol. 41, No. 3. P. 285−311.

Frolova E. N. Gaponov S. P. Eastern steppe viper Vipera renardi (Christoph, 1861) in the Divnogorye museum-reserve, Trudy Karel'skogo nauchnogo centra RAN. 2017. No. 7. P. 93−99.

Gagina T. N. Skalon V. N. Reptiles of Eastern Siberia, Gerpetologiya. Tashkent: Nauka, 1965. P. 17−23.

Garanin V. I. Pavlov A. V. Bakiev A. G. Steppe viper, or Renard's viper Vipera renardi (Christoph, 1861), Zmei Volzhsko-Kamskogo kraya. Samara: Izd-vo Samarskogo nauchnogo centra RAN, 2004. C. 61−90.

Garanin V. I. Amphibians and reptiles of the Volga-Kama region. M.: Nauka, 1983. 176 p.

Giovanelli G. R., Haddad C. F. B., Alexandrino J. Predicting the potential distribution of the alien invasive American bullfrog (Lithobates catesbeianus) in Brasil, Biol. Invasions. 2007. Vol. 10, No. 5. P. 585−590.

Gloyd H. K., Conant R. Snakes of the Agkistrodon Somplex. A monographic review, SSAR. Contributions to Herpetology. No. 6. Oxford, Ohio, 1990. 614 p.

Golovcov D. E. Vertebrates of the Chatkal Reserve, Trudy Chatkal'skogo gosudarstvennogo biosfernogo zapovednika. 2007. No. 6. P. 178−220.

Golubev M. L. New finds of amphibians and reptiles in Kazakhstan, Vestnik zoologii. 1990. No. 5. P. 76−78.

Golubev M. L. Phrynocephalus guttatus (Gmel.) or Ph. versicolor Str. (Reptilia, Agamidae): what kind of toadhead agama lives in Kazakhstan?, Vestnik zoologii. 1989. No. 5. P. 38−46.

Grazhdankin A. V. Reactions of reptiles of arid territories to high temperatures and insolation, Zoologicheskiy zhurnal. 1973. T. 52, No. 4. P. 552−560.

Groff L. A., Marks Sh. B., Hayes M. P. Using Ecological Niche Models to Direct Rare Amphibian Surveys: A Case Study Using the Oregon Spotted Frog (Rana pretiosa), Herpetological Conservation and Biology. 2014. Vol. 9, No. 2. P. 354−368.

Gvoždík V., Jandzik D., Cordos B., Rehák I., Kotlík P. A mitochondrial DNA phylogeny of the endangered vipers of the Vipera ursinii complex, Molecular Phylogenetics and Evolution. 2012. No. 62. P. 1019−1024.

Hamitov A. Zh. Species diversity, expansion, distribution of snakes in the forest-steppe and steppe zones of Kazakhstan, Principy ekologii. 2016. T. 5, No. 3. P. 154−155.

Hinkle D. E., Wiersma W., Jurs S. G. Applied Statistics for the Behavioral Sciences. 5th ed. Boston: Houghton Mifflin, 2003. 756 p.

Ishunin G. I. Poisonous snakes of Uzbekistan. Tashkent: Izd-vo AN UzSSR, 1950. 31 p.

Jueterbock A., Smolina I., Coyer J. A., Hoarau G. The fate of the Arctic seaweed Fucus distichus under climate change: an ecological niche modeling approach, Ecology and Evolution. 2016. Vol. 6, No. 6. P. 1712−1724.

Karpenko V. P. Distribution and ecology of the Halys pit viper Ancistrodon halys (Pallas, 1779): Abstract of the dissertation for the degree of Candidate of Biological Sciences. Tashkent: Izd-vo AN UzSSR, 1958. 19 p.

Karpenko V. P. On the ecology of the steppe viper Vipera ursini renardi (Christoph) in the spring-summer season, Yadovitye zhivotnye Sredney Azii i ih yady: Materialy Sredneaziatskoy konf., 1−3 oktyabrya 1968 g. Tashkent, 1970. P. 77−87.

Kaschenko N. F. Results of the Altai zoological expedition in 1898. Vertebrates. With tables 1-4. Tomsk: Tipo-litografiya M. N. Kononova i I. F. Skulimovskogo, 1899. 158 p.

Kaschenko N. O. Reptiles collected by the Central Asian expeditions of prof. V.V. Sapozhnikov in 1902-1906 and 1908, Ezhegodnik Zoologicheskogo muzeya Imperatorskoy Akademii nauk. SPb., 1909. P. 119−130.

Kashkarov D. N. Ecological sketch of the region of lakes: Biili-kul, Ak-kul, and Ashchi-kul of the Aulieata district. Tashkent: Izd-vo Sredn, Aziat. gop. un-ta, 1928. 55 p. (Trudy Sredne-Aziatskogo gosudarstvennogo universiteta. Seriya 8-a. Zoologiya. No. 2).

Kashkarov D. N. Zooecological sketch of the eastern part of the Betpak-Dala desert. Tashkent: Izd-vo Sredneaziatskogo gop. un-ta, 1935. 30 p.

Kassal B. Yu. Common viper Vipera berus (Linnaeus, 1758), Krasnaya kniga Omskoy oblasti. Omsk: Izd-vo OmGPU, 2006. P. 71−72.

Kireev V. A. Amphibians and reptiles of the Zheltau ridge, Voprosy gerpetologii. L.: Nauka, 1981. P. 64−65.

Kolbincev V. G. Fauna and distribution of amphibians and reptiles of Small Karatau and adjacent territories, Gerpetologicheskie issledovaniya v Kazahstane i sopredel'nyh stranah. Almaty, 2010. P. 100−117.

Kolobaev N. N. On the influence of microclimate on the morphological features of the Halys pit viper, Vliyanie izmeneniya klimata na ekosistemy basseyna reki Amur. M.: WWF Rossii, 2006. P. 120−128.

Kostin V. P. Notes on the distribution and ecology of amphibians and reptiles of the ancient delta of the Amu Darya and Kara-Kalpak Ustyurt, Trudy Instituta zoologii i parazitologii AN Uzb SSR. T. 5. Zoologicheskiy sbornik. Tashkent, 1956. P. 47−66.

Kotlov A. A. Species composition and distribution of amphibians and reptiles of southwestern Kulunda, Altayskiy zoologicheskiy zhurnal. 2008. No. 2. P. 131−134.

Kovalenko V. G. Steppe viper: Avtoref. dip. ... kand. biol. nauk. Alma-Ata, 1952. 12 p.

Kren' A. K. Materials on the fauna of reptiles and mammals of the Arys-Kum desert, Uchenye zapiski Alma-Atinskogo gosudarstvennogo pedagogicheskogo i uchitel'skogo institutov. 1953. T. 3, No. 2. P. 134−150.

Kropachev I. I. Orlov N. L. A new subspecies of the Halys pit viper Gloydius halys (Pallas, 1776) (Viperidae, Crotalinae) from Tuva and western Mongolia, Trudy Zoologicheskogo instituta RAN. 2017. T. 321, No. 2. P. 129−179.

Kropachev I. I. Analysis of the geographical and landscape distribution of amphibians and reptiles in the Republic of Tuva, Trudy Zoologicheskogo instituta RAN. 2014. T. 318, No. 4. P. 486−562.

Kubykin R. A. Ecological and faunistic review of the reptiles of the islands of Alakol lake, Izvestiya AN KazSSR. 1975. No. 3. P. 10−16.

Kubykin R. A. The amazing world of reptiles. M.: Nauchnyy mir, 2003. 264 p.

Kubykin R. A., Brushko Z. K. Contemporary spreading and information on Agkistrogon halys caraganus Eichwald, 1831 (Reptilia, Crotalidae) numbers in Kazakhstan, Bectnik KazGU. Ser. biologicheskaya. 1998. No. 6. P. 9−13.

Kuranova V. N. Kolbincev V. G. Biology of the common adder (Vipera berus) in the Tomsk region, Voprosy gerpetologii. L.: Nauka, 1981. P. 80−81.

Kuranova V. N. Simonov E. P. Yarcev V. V. Shamgunova R. R. Starikov V. P. Diversity, distribution and conservation status of reptiles in Western Siberia, Gerpetologicheskie issledovaniya v Kazahstane i sopredel'nyh stranah. Almaty, 2010. 149 p.

Kuranova V. N. Zinchenko V. K. Population differences in bioproductivity indicators of the common adder in the South-East of Western Siberia, Voprosy gerpetologii. L.: Nauka, 1985. P. 118−119.

Lada G. A. Moulting and seasonal feeding activity of the common adder in the Tambov region, Voprosy gerpetologii. L.: Nauka, 1981. P. 82.

Le Neve A., Gouraud C., Morion F., Judas J. Kazakhstan Nature Trip Report 2009. National Avian Research Center, ResearchGate. 2010. URL: https://www.researchgate.net/publication/309533797_Kazakhstan_Nature_Trip_Report_2009 (data obrascheniya: 12.05.2019).

Lesnyak A. P. Confinement to shelters and daily activity of the western Halys pit viper in South Kazakhstan, Pozvonochnye zhivotnye Sredney Azii. Tashkent: Fan, 1966. P. 187−193.

Lisovskiy A. A. Obolenskaya E. V. A study of the distribution ranges of small mammals from South-Eastern Transbaikalia with ecological niche modeling methods, Zhurnal obschey biologii. 2014. T. 75, No. 5. P. 353−371.

Liu C., White M., Newell G. Selecting thresholds for the prediction of species occurrence with presence-only data, Journal of Biogeography. 2013. Vol. 40, No. 4. P. 778−789.

Makarova T. N. Morphological characteristics of newborn Renard's vipers Vipera renardi from the Lower Volga region, Samarskaya Luka: problemy regional'noy i global'noy ekologii. 2013. T. 22, No. 4. P. 36−39.

Malmanov B. D. Report on the implementation of scientific research for 1987 on the topic: Herpetofauna of the Western chin of Ustyurt, the Karynzharyk depression and adjacent areas. Novyy-Uzen', 1987. 10 p.

Markov G. S. Bogdanov O. P. Fomina M. I. Chernobay V. F. Parasite fauna of the steppe viper − Vipera ursini (Bonaparte) in Kazakhstan, Voprosy gerpetologii i toksikologii zmeinyh yadov. Tashkent: Nauka, 1966. P. 22−26.

Merow C., Smith M. J., Silander J. A. Jr. A practical guide to MaxEnt for modeling species’ distributions: what it does, and why inputs and settings matter, Ecography. 2013. No. 36. P. 1058−1069.

Mil'ko D. A. Panfilov A. M. Amphibians and reptiles, Krasnaya kniga Kyrgyzskoy Respubliki. 2-e izd. Bishkek: GAOOSiLH, BPI NAN KR, EDK «Aleyne», 2007. P. 315−345.

Nakarenok E. G. Ecological aspects of the formation of the herpetofauna of the northern Caspian and the trends of its modern development: Dissertations for the degree of candidate of biological sciences. N. Novgorod, 2002. 206 p.

Nekrasova O. D. Titar V. M. Modeling and bioclimatic analysis of changes in the range of dice snake Natrix tessellata (Reptilia, Colubridae) in Ukraine, Pracі ukraїns'kogo gerpetologіchnogo tovaristva. 2014. No. 5. P. 80−83.

Nikol'skiy A. M. Fauna of Russia and adjacent countries. Reptiles (Reptilia). Vol. 2. Ophidia. Pg.: Imperatorskaya akademiya nauk, 1916. 350 p.

Nikol'skiy A. M. On the vertebrate fauna of the bottom of the Balkhash Basin. SPb., 1887. 130 p.

Nikol'skiy A. M. Reptiles and amphibians collected by P.P. Sushkin in the Turgai region, Bull. Natur. de Moscou. 1899. No. 4. P. 366−368.

Nilson G., Andren C. The Meadow and Steppe vipers of Europe and Asia − The Vipera (Acridophaga) ursinii complex, Acta Zoologica Academiae Scientiarum Hungaricae. 2001. Vol. 47, No. 2-3. P. 87−267.

Nuridzhanov D. A. Abduraupov T. V. Gricyna M. A. Results of a herpetological survey of the southern and central parts of the Ustyurt plateau and the Sarykamysh basin (Republic of Uzbekistan), Vestnik Tyumenskogo gosudarstvennogo universiteta. Ekologiya i prirodopol'zovanie. 2019. T. 5, No. 1. P. 74−99.

Nuridzhanov D. A. The current state of the abundance and distribution of reptiles on the Ustyurt Plateau (Uzbekistan), Nazemnye pozvonochnye zhivotnye aridnyh ekosistem: Materialy Mezhdunar. konf., posvyasch. pamyati N. A. Zarudnogo (Tashkent, 24−27 oktyabrya 2012 g.). Tashkent, 2012. P. 248−252.

Ogurcov S. S. Modeling of habitat suitability and distribution of brown bear (Ursus arctos) in the Southern taiga subzone using the method of maximum entropy, Nature Conservation Research. Zapovednaya nauka. 2019. Vol. 4, Issue 4. P. 34−64.

Okulova N. M. K biologii stepnoy gadyuki (Vipera ursini) v Zapadnom Kazahstane [On the biology of the steppe viper (Vipera ursini) in Western Kazakhstan, Voprosy gerpetologii. L.: Nauka, 1981. P. 93−94.

Orlov N. L., Barabanov A. V. Analysis of Nomenclature, Classification, and Distribution of the Agkistrodon halys – Agkistrodon intermedius Complexes: a Critical Review, Russian Journal of Herpetology. 1999. Vol. 6, No. 3. P. 167−192.

Orlova V. F. Terbish H. Materials on the herpetofauna of the Dzungarian Gobi, Gerpetologicheskie issledovaniya v Mongol'skoy Narodnoy Respublike: Sbornik nauchnyh trudov. M., 1986. P. 95−110.

Pallas P. S. Bemerkungen auf einer Reise in die sudlichen statthalterschaften des Russischen Reichs in den jahren 1793 und 1794. Erster Band. Leipzig: G. Martini, 1799. 516 p.

Pallas P. S. Travelling to different provinces of the Russian Empire. Part 1. SPb., 1773. 658 p.

Paporotnyy D. I. On the biology of Halys pit viper in the conditions of Barsa-Kelmes Island, Trudy gosudarstvennogo zapovednika Barsa-Kel'mep. Vyp. 1. Alma-Ata, 1950. P. 136−146.

Paraskiv K. P. Butovskiy P. M. About the fauna of amphibians and reptiles of Western Kazakhstan, Trudy Instituta zoologii AN Kaz. SSR. 1960. T. 13. P. 148−159.

Paraskiv K. P. Reptiles of Kazakhstan. Alma-Ata: Izd-vo AN KazSSR, 1956. 228 p.

Pavlov A. V. Petrova I. V. Karmazina I. O. Orthoptera (Ortoptera) in the diet of the eastern steppe viper (Vipera renardi) at the northern limit of the range, Voprosy gerpetologii. SPb., 2011. P. 208−212.

Pavlov A. V. Zinenko A. I. Yoger U. Shtumpel' N. Petrova I. V. Malenev A. L. Zayceva O. V. Shurshina I. V. Bakiev A. G. Natural hybridization of eastern steppe vipers Vipera renardi and common adder V. berus, Izvestiya Samarskogo nauchnogo centra Rossiyskoy akademii nauk. 2011. T. 13, No. 5. P. 172−178.

Pestinskiy B. V. Materials on the biology of poisonous snakes in Central Asia, their catching and keeping in captivity, Trudy Uzbekistanskogo Zoologicheskogo sada. T. 1. Tashkent: Gostehizdat UzSSR, 1939. P. 38−53.

Pestov M. V. Saraev F. A. Ageev V. S. New finds of reptiles in the Northern Caspian (Republic of Kazakhstan), Sovremennaya gerpetologiya. 2011. T. 11, No. 3/4. P. 192−195.

Pestov M. V. Saraev F. A. Findings of nests of some raptors on the Akkeregeshin Cretaceous plateau, Atyrau region, Kazakhstan, Pernatye hischniki i ih ohrana. 2009. No. 15. P. 132−133.

Pestov M. V. Halys pit viper − a new species of fauna of the Novosibirsk region, Zemnovodnye i presmykayuschiesya Novosibirskoy i Tomskoy oblastey: Inform. materialy k gerpetofaune Sibiri. Novosibirsk: Izd-vo Novosib. gop. ped. un-ta, 2003. P. 35−38.

Petrov V. Yu. Krymov N. G. Steppe viper − Vipera ursini (Bonaparte, 1835), Krasnaya kniga Altayskogo kraya. Redkie i nahodyaschiesya pod ugrozoy ischeznoveniya vidy zhivotnyh. T. 2. Barnaul: Izd-vo Altayskogo un-ta, 2016. P. 108−110.

Petrova I. V. Pavlov A. V. Chizhikova N. A. Naumkina N. A. Status of common adder (Vipera berus L.) populations in PAs depending on indicators of anthropogenic load, Principy i sposoby sohraneniya bioraznoobraziya: Materialy III Vserop. nauch. konf. Yoshkar-Ola; Puschino: Mar. gop. un-t, 2008. P. 570−573.

Phillips S. J., Anderson R. P., Schapire R. E. Maximum entropy modeling of species geographic distributions, Ecological Modelling. 2006. Vol. 190. P. 231−259.

Phillips S. J., Dudik M. Modeling of species distributions with Maxent: new extensions and a comprehensive evaluation, Ecography. 2008. Vol. 31. P. 161−175.

Phillips S. J., Dudik M., Schapire R. E. A Maximum Entropy Approach to Species Distribution Modeling, Proceedings of the Twenty-First International Conference on Machine Learning. Banff, 2004. P. 655−662.

Pomazenko O. A. Tabachishin V. G. Morphogenetic characteristics of populations of the eastern steppe viper Vipera renardi in the lower Volga region and adjacent territories, Izvestiya Saratovskogo universiteta. Nov. ser. Ser. Himiya. Biologiya. Ekologiya. 2017. T. 14, No. 4. P. 104−109.

Popov V. A. Lukin A. B. Animal world of Tataria. (Vertebrates). Kazan': Tatgosizdat, 1949. 218 p.

Popudina A. D. Features of the lifestyle of the common adder in the south of the Ob forest zone, Fauna i ekologiya zhivotnyh Priob'ya. Vyp. 133. Novosibirsk, 1976. P. 51−54.

Prokopov K. P. Herpetofauna of the Katon-Karagai State National Natural Park, Trudy Katon-Karagayskogo gosudarstvennogo nacional'nogo prirodnogo parka. T. 1. Ust'-Kamenogorsk: Profit, 2006. P. 131−147.

Radzhabov B. Yadgarov T. Davlyatov Ya. D. Some rare and endangered snake species of Uzbekistan, Ohrana zhivotnogo mira i rasteniy Uzbekistana: Tez. dokl. konf. Tashkent: Fan, 1978. P. 32−33.

Ruzskiy M. D. Zoodynamics of the Baraba steppe, Trudy Tomskogo universiteta. 1946. No. 97. P. 17−68.

Saraev F. A. Pestov M. V. To the cadastre of reptiles of the Northern and North-Eastern Caspian, Gerpetologicheskie issledovaniya v Kazahstane i sopredel'nyh stranah. Almaty, 2010. P. 172−191.

Selevin V. A. Faunistic materials, Trudy Sredneaziatskogo gosudarstvennogo universiteta. Seriya 8-a. Zoologiya. 1935. No. 21. P. 20−22.

Semenov D. V. Shenbrot G. I. Materials on the herpetofauna of southeast Mongolia, Gerpetologicheskie issledovaniya v Mongol'skoy Narodnoy Respublike: Sb. nauch. tr. M., 1986. P. 110−119.

Shammakov S. Reptiles of lowland Turkmenistan, AN Turkmenskoy SSR. Institut zoologii. Ashhabad: Ylym, 1981. 311 p.

Shebzuhova E. A. Reptiles of the steppe zone of the Adygei Autonomous Region, Voprosy gerpetologii. L.: Nauka, 1981. P. 155−156.

Shi J. Yang D. Zhang W. Li P. Ding L. In Chinese

Shilov M. N. A case of carrion eating by Halys pit viper, Byulleten' MOIP. Otd. biol. 1956. T. 61, No. 4. Iyul'-Avgust. P. 86.

Shilov M. N. Materials on terrestrial vertebrates of Kazakhstan, Trudy Instituta zoologii AN KazSSr. 1961. T. 15. P. 170−176.

Shilov M. N. Notes on some reptiles of the Northern Aral Sea, Trudy Instituta zoologii AN KazSSR. 1961. T. 15. P. 170−177.

Shilova E. S. About wintering of the Halys pit viper, Byulleten' MOIP. Otd. biol. 1956. T. 61. No. 4. Iyul'-Avgust. P. 86−87.

Shnitnikov V. I. Reptiles of Semirechye, Trudy obschestva izucheniya Kazahstana. T. 8, No. 3. Kzyl-Orda: Izdanie obschestva izucheniya Kazahstana, 1928. 85 p.

Shul'pin L. M. Materials on mammals and reptiles of the Talas Alatau, Izvestiya AN KazSSR. Ser. biol. 1948. T. 51, No. 7. P. 65−83.

Shutova Yu. V. Class Reptiles Reptilia, Krasnaya kniga Altayskogo kraya. Redkie i nahodyaschiesya pod ugrozoy ischeznoveniya vidy zhivotnyh. T. 2. Barnaul: OAO «IPP «Altay», 2006. P. 43−46.

Sidorov S. A. Amphibians and reptiles of the Aral Sea, Byulleten' MOIP. Otd. biol. 1925. T. 33, No. 1-2. P. 188−200.

Sillero N., Campos J., Bonardi A., Corti C., Creemers R., Crochet P., Isailović J. C., Denoël M., Ficetola G. F., Gonçalves J., Kuzmin S., Lymberakis P., de Pous Ph., Rodríguez A., Sindaco R., Speybroeck J., Toxopeus B., Vieites D. R., Vences M. Updated distribution and biogeography of amphibians and reptiles of Europe, Amphibia-Reptilia. 2014. Vol. 35, No. 1. P. 1−31.

Simonov E. P. Clarification of the northern border of the range of the Halys pit viper (Gloydius (Agkistrodon) halys) and its biotope confinement in the territory of Western Siberia, Zoologiya. Ser. biol. 2008. P. 65−70.

Sindaco R., Venchi A., Grieco C. The Reptiles of the Western Palearctic. 2. Annotated checklist and distributional atlas of the snakes of Europe, North Africa, Middle East and Central Asia, with an update to the Vol. 1. Letina (Italy): Edizioni Belvedere, 2013. 544 p.

Smirin V. M. On the nesting of the short-toed snake-eagle in the Aral, Zoologicheskiy zhurnal. 1959. T. 38, No. 11. P. 1756−1757.

Smirnovskiy B. N. Poisonous snakes of Kazakhstan and their significance as pests in animal husbandry: Abstract of the dissertation of the candidate of biological sciences. Alma-Ata, 1961. 15 p.

Smirnovskiy B. N. Poisonous snakes of Kazakhstan. Alma-Ata: Kazsel'hozgiz, 1963. 64 p.

Sokolov A. S. On the variability of the pylaeus of the common adder (Vipera berus) of the Oka-Don Plain, Voprosy gerpetologii. Puschino; M., 2001. P. 270−272.

Stohlgren T. J., Jarnevich C. S., Esaias W. E., Morisette J. T. Bounding species distribution models, Current Zoology. 2011. Vol. 57, No. 5. P. 642−647.

Tabachishin V. G., Yermokhin M. V. Distribution of Vipera renardi renardi (Christoph, 1861), in the Saratov region, Russian Federation, Herpetozoa. 2019. Vol. 31, No. 3-4. P. 233−235.

Tabachishina I. E. Tabachishin V. G. Shlyahtin G. V. To clarify the northern border of the distribution of the eastern steppe viper (Vipera renardi) in the Volga region, Povolzhskiy ekologicheskiy zhurnal. 2007. No. 3. P. 271−277

Tabachishina I. E. Tabachishin V. G. Zav'yalov E. V. Morphoecological characteristics of the Lower Volga populations of the steppe viper (Vipera ursini), Povolzhskiy ekologicheskiy zhurnal. 2002. No. 1. P. 76−81.

Terent'ev P. V. Chernov S. A. Key to reptiles and amphibians. M.: Sov. nauka, 1949. 340 p.

Tkachenko S. V. Verzhuckiy D. B. Holin A. V. Discovery of the Halys pit viper (Agkistrodon halys Pall., 1775) in Southwestern Tuva, Baykal'skiy zoologicheskiy zhurnal. 2010. T. 2, No. 5. P. 34−35.

Toporkova L. Ya. Amphibians and reptiles of the Urals, Fauna evropeyskogo Severa, Urala i Zapadnoy Sibiri. Sverdlovsk: Ural'skiy rabochiy, 1973. P. 84−117.

Tupikov A. I. Ukrainskiy P. A. Comparative analysis of different approaches to modeling the species range in the Maxent program (on the example of the patterned snake and the steppe viper), Nauchnye vedomosti Belgorodskogo gosuniversiteta. Seriya Estestvennye nauki. 2016. T. 4 (225), No. 34. P. 71−84.

Tupikova N. V. Komarova L. V. Principles and methods of zoological mapping. M.: Izd-vo MGU, 1979. 192 p.

Vakker V. G. Parasite fauna of reptiles in the south of Kazakhstan and their role in the circulation of some human and animal helminths: Dip. … kand. biol. nauk. Alma-Ata: Institut zoologii AN KazSSR, 1971. 365 p.

Vashetko E. V. Chikin Yu. A. Hodzhaev A. F. Nuridzhanov A. S. Herpetofauna of the Western Tien Shan, Sovremennaya gerpetologiya. 2003. T. 2. C. 24−38.

Vedmederya V. I. Zinenko A. I. Goncharenko L. A. Catalog of the collections of the Museum of Nature of V. N. Karazin Kharkiv National University. Serpents (Reptilia: Serpentes). Har'kov, 2007. 82 p.

Velikanov V. P. About the herpetofauna of the Sarykamysh depression, Voprosy gerpetologii. L.: Nauka, 1977. P. 56−57.

Vershinin V. L. Amphibians and reptiles of the Urals. Ekaterinburg: UrO RAN, 2007. 227 p.

Vozniychuk O. P. Kuranova V. N. Amphibians and reptiles of the Katunsky Reserve and adjacent territory (Central Altai), Sovremennaya gerpetologiya. 2008. T. 8, No. 2. P. 101−117.

Wang T., Wang G., Innes J., Nitschke C., Kang H. Climatic niche models and their consensus projections for future climates for four major forest tree species in the Asia–Pacific region, Forest Ecology and Management. 2016. No. 360. P. 357−366.

Yakovlev V. A. Cadastre of amphibians and reptiles of the Republic of Altai, Zhivotnyy mir Altae-Sayanskoy gornoy strany: Sb. nauch. tr. Gorno-Altaysk: RIO «Univer-Print» GAGU, 1999. P. 175−214.

Yakovlev V. A. On the issue of the eastern border of the range of the steppe viper, Biologicheskie resursy Altayskogo kraya i perspektivy ih ispol'zovaniya: Tez. dokl. k konf. Barnaul, 1984. P. 41.

Yakovlev V. A. Steppe viper − Vipera ursini (Bonaparte, 1835), Krasnaya kniga Respubliki Altay (zhivotnye, 3-e izd.). Gorno-Altaysk, 2017. P. 117−118.

Yakovleva I. D. Reptiles of Kyrgyzstan. Frunze: Izd-vo AN Kirg. SSR, 1964. 273 p.

Zav'yalov E. V. Tabachishin V. G. Shlyahtin G. V. Kaybeleva E. I. Mosolova E. Yu. Tabachishina I. E. Yakushev N. N. Cataloging of zoological collections. Issue 2. Stock collections in the herpetofauna monitoring system. Saratov: Izd-vo Sarat. un-ta, 2006. 96 p.

Zhao E. M., Huang M. H., Zhong Y. Fauna Sinica: Reptilia. Vol. 3. Squamata, Serpentes. Beijing: Science Press, 1999. 460 p.

Zhdokova M. K. Distribution and some aspects of morphology of the steppe viper Vipera ursinii in Kalmykia, Sovremennaya gerpetologiya. 2003a. T. 2. P. 143−147.

Zhdokova M. K. Ecological and morphological analysis of the amphibian and reptile fauna of Kalmykia: Dip. … kand. biol. nauk. Saratov, 2003b. 262 p.

Zima Yu. A. The current state of the population of the steppe viper Vipera renardi (Christoph, 1861) in the foothills of the Trans-Ili Alatau (Almaty region), Zoologicheskie issledovaniya za 20 let nezavisimosti Respubliki Kazahstan: Materialy Mezhdunar. nauch. konf. Almaty, 2011. C. 241−242.

Zinchenko V. K. Zinchenko Yu. K. Distribution of amphibians and reptiles of the Markakol state reserve, Ohrana okruzhayuschey sredy i prirodopol'zovanie Priirtysh'ya. Ch. 2. Ust'-Kamenogorsk, 1990. P. 141−143.

Zinenko O., Stümpel N., Mazanaeva L. F., Bakiev A., Shiryaev K., Pavlov A., Kotenko T., Kukushkin O., Chikin Y., Duisebayeva T., Nilson G., Orlov N. L., Tuniyev S. B., Ananjeva N. B., Murphy R. W., Joger U. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi), Molecular Phylogenetics and Evolution. 2015. No. 84. P. 85−100.